In this system, two cycles are connected through a joint catalyzing kinase [Feliu, Wiuf].
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In the motif Modification of Two Different Substrates H, the symbols used in the chemical reactions have the following meaning:
S_i and P_i are substrates with i = 0, 1, 2 phosphorylated sites, E is a kinase and F is a phosphotase, X and Y are the intermediate complexes formed by the enzymes E and F, respectively.
To rename species, use substitute(ReactionNetwork,List).
The object modificationOfTwoSubstratesH is a method function.